North Rona lies on the outer reaches of the Outer Hebrides, one of the remotest islands in Europe. It lost its last significant population in the late 1600s, the island’s inhabitants dying after a colony of black rats swarmed ashore from a shipwreck. The rodents either ate the islanders’ food stores or spread the plague. Either way, it would have been a bleak end.

The island was largely used as pasture in the centuries after, by folk such as the King. Now owned by Scottish Natural Heritage, its inhabitants are a little less woolly ­- and a lot more colourful.

We were in the Outer Hebrides to survey the remote caves of St Kilda and North Rona and document the wildlife that called them home. A small number of biological records do exist regarding the sea caves around St Kilda, but there is nothing published about those of North Rona. Prior to our visit, no formal survey work had been conducted at either site.

In 1992, as a result of the Biodiversity Convention at the Rio Earth Summit, the European Community implemented the Habitats Directive. This is the foundation of Europe’s nature conservation policy and was formed to protect its most valuable and threatened species and habitats. Central to the Habitats Directive is the Natura 2000 network, a series of protected marine and terrestrial areas stretching right across Europe. One of the classifications for protected sites is the Special Area of Conservation (SAC). SAC status is assigned to specific habitats and associated wildlife listed under Annex I and II of the Habitats Directive.